Joe "The Boss" Masseria (?-1931)- This guy was the man in NY until he decided to piss off the wrong people. He is responsible for starting the Castellammarese War and died because of it in 1931. His death signified the end of the war.

Charles "Lucky" Luciano (1931-1946)- Charlie Lucky replaced Masseria as the head of the Masseria family in 1931, after the end of the war. He was one of the seven original members of the Commission. He became the main power when Salvatore Maranzano was murdered in 1931. He stayed in power until the mid-40s when he was deported.

Frank Costello (1946-1957)- Was known as the "Prime Minister" because he would much rather work things out than fight over them. He was one of Cosa Nostra's true earners and was known to be the best political corrupter ever. His reign as boss ended in 1957 after an attempt on his life.

Vito Genovese (1957-1969)- After the retirement of Costello in 1957, he became boss. He had been a power in the families for years but had never been boss. Now he took it away from Costello and ran right into the hands of authorities. Genovese was the man who set up Appalachin in 1957 and ended up losing much power after that disaster. Soon he was convicted and ruled from prison until 1969 when he died.

"Tommy Ryan" Eboli (1969-1972)- Man who took over after the death of Vito Genovese. However, only a few years later, he was murdered (in 1972).

Anthony "Fat Tony" Salerno (1972-1987)- Soon after the death of "Tommy Ryan", Tony ended up a very powerful boss. He was in power during most of the 70s and was finally apprehended by the authorities in 1987 during the "Commission Trials". He died in prison on July 29, 1992.

Vincent "The Chin" Gigante (1987-1998)- Took over power after "Fat Tony" went up the river in 1987. Most people would have never suspected the assassin who tried to kill Frank Costello three decades earlier would have ever risen to power. He was the most powerful man in NY but his insanity ploy finally caught up with him and he was found guilty of racketeering and murder conspiracy on July 25, 1997. He was senteced to 12 years in prison and $1.25 million in fines.

Dominick "Quiet Dom" Cirillo (1998-present)- Just became the official boss after "Oddfather" got 12. "Quiet Dom" Cirillo, who as opposed to Gigante and his bathrobe, prefers slacks and a tee-shirt. He's not bringing a lot of attention to himself and is the most powerful boss in the country, with his family of about 250-300 members, by far the largest, with the Gambino's being the only family coming close with about 200 members.